List of Hypericum species

Hypericum perforatum, the type species of the genus Hypericum^[1]

The genus Hypericum contains the following 490 species, according to The Plant List,^[2] divided by sections.^[3] Hypericum species are found most densely in Europe and parts of the Middle East.^[4] They are a rather unusual genus in the fact that Norman Robson created a genus-wide monograph in 1977, dividing the genus into distinct sections that are still used today despite the genus's size ^[5] A phylogenetic study was completed for Hypericum, showing that the genus Triadenum is likely a clade in Hypericum. It also showed that Thornea is sister to Hypericum and that 60% of the sections of Hypericum are monophyletic.^[6]

Adenosepalum

Adenosepalum Spach contains shrubs, shrublets, and soft herbs. Its species grow to 2.5 meters tall and can be found in most of Europe and parts of Northern Africa.

Hypericum abilianum N.Robson
Hypericum aciculare Kunth
Hypericum afromontanum Bullock
Hypericum kiboense Oliv.
Hypericum lanuginosum Lam.
Hypericum montanum L.
Hypericum naudinianum Coss. & Durieu
Hypericum pubescens Boiss.
Hypericum sinaicum Hochst. ex Boiss.
Hypericum tomentosum L.

Adenotrias

Adenotrias (Jaub. & Spach) R. Keller contains shrubs and shrublets. Its species grow up to 2 meters tall and are found in West Africa and Eastern Europe.

Hypericum aciferum (Greuter) N. Robson
Hypericum aegypticum L.
Hypericum russeggeri (Fenzl) R. Keller

Androsaemum

The species in Androsaemum (Duhamel) Godron grow up to 3 meters tall and are shrubs. They are found in the Middle East and around the Mediterranean.

Hypericum androsaemum L.
Hypericum foliosum Aiton
Hypericum grandifolium Choisy
Hypericum hircinum L.

Arthrophyllum

The species in Arthrophyllum Jaub. & Spach grow to be less than a meter tall. They are found in parts of the Middle East.

Hypericum cardiophyllum Boiss.
Hypericum nanum Poir.
Hypericum pamphylicum N. Robson & P.H.Davis
Hypericum rupestre Jaub. & Spach
Hypericum vacciniifolium Hayek & Siehe

Ascyreia

Ascyreia Choisy contains shrubs, shrublets, and rarely trees. Most species grow to be 4–5 m tall and can be found across Europe.

Hypericum acmosepalum N.Robson
Hypericum augustinii N.Robson
Hypericum beanii N.Robson
Hypericum bellum H.L.Li
Hypericum calycinum L.
Hypericum choisianum Wall. ex N.Robson
Hypericum cohaerens N.Robson
Hypericum cordifolium Choisy
Hypericum curvisepalum N.Robson
Hypericum gaitii Haines
Hypericum forrestii (Chitt.) N.Robson
Hypericum gracilipes Stapf ex C.E.C.Fisch.
Hypericum griffithii Hook.f. & Thomson ex Dyer
Hypericum henryi H.Lév. & Vaniot
Hypericum hookerianum Wight & Arn.
Hypericum kouytchense H.Lév.
Hypericum lacei N.Robson
Hypericum lagarocladum N.Robson
Hypericum lancasteri N.Robson
Hypericum leschenaultii Choisy
Hypericum lobbii N.Robson
Hypericum maclarenii N.Robson
Hypericum monogynum L.
Hypericum mysurense Wall. ex Wight & Arn.
Hypericum oblongifolium Choisy
Hypericum pachyphyllum Collett & Hemsl.
Hypericum patulum Thunb.
Hypericum podocarpoides N.Robson
Hypericum prattii Hemsl.
Hypericum pseudohenryi N.Robson
Hypericum reptans Hook.f. & Thomson ex Dyer
Hypericum sherriffii N.Robson & D.G.Long
Hypericum siamense N.Robson
Hypericum stellatum N.Robson
Hypericum subsessile N.Robson
Hypericum tenuicaule Hook.f. & Thomson ex Dyer
Hypericum uralum Buch.-Ham. ex D.Don
Hypericum wardianum N. Robson
Hypericum williamsii N.Robson
Hypericum wilsonii N.Robson

Brathys

Brathys (Mutis ex L.f.) Choisy is one of the largest sections of Hypericum. It contains mainly evergreen trees, shrubs, shrublets, and wiry herbs that grow up to 6 m tall.

Hypericum acostanum Steyerm. ex N.Robson
Hypericum addingtonii N.Robson
Hypericum andinum Gleason
Hypericum arbuscula Standl. & Steyerm.
Hypericum asplundii N. Robson
Hypericum baccharoides Cuatrec.
Hypericum beamanii N. Robson
Hypericum bolivaricum N. Robson
Hypericum brathys Sm.
Hypericum bryoides Gleason
Hypericum callacallanum N. Robson
Hypericum caracasanum Willd.
Hypericum cardonae Cuatrec.
Hypericum carinosum R.Keller
Hypericum cassiopiforme N. Robson
Hypericum castellanoi N.Robson
Hypericum chamaemyrtus Triana & Planch.
Hypericum costaricense N.Robson
Hypericum cuatrecasii Gleason
Hypericum cymobrathys N. Robson
Hypericum decandrum Turcz.
Hypericum diosmoides Griseb.
Hypericum drummondii (Grev. & Hook.) Torr. & A. Gray
Hypericum eastwoodianum I.M.Johnst.
Hypericum elatoides R.Keller
Hypericum espinalii N. Robson
Hypericum fuertesii Urb.
Hypericum galinum S.F. Blake
Hypericum garciae Pierce
Hypericum gentianoides (L.) Britton, Sterns & Poggenb.
Hypericum gladiatum N. Robson
Hypericum gleasonii N. Robson
Hypericum gnidioides Seem.
Hypericum goyanesii Cuatrec.
Hypericum harlingii N. Robson
Hypericum hartwegii Benth.
Hypericum horizontale N. Robson
Hypericum humboldtianum Steud.
Hypericum irazuense Kuntze ex N.Robson
Hypericum jaramilloi N.Robson
Hypericum juniperinum Kunth
Hypericum lancifolium Gleason
Hypericum lancioides Cuatrec.
Hypericum laricifolium Juss.
Hypericum laricoides Gleason
Hypericum llanganaticum N. Robson
Hypericum loxense Benth.
Hypericum lycopodioides Triana & Planch.
Hypericum magdalenicum N.Robson
Hypericum magniflorum Cuatrec.
Hypericum maguirei N. Robson
Hypericum marahuacanum N.Robson
Hypericum martense N. Robson
Hypericum matangense N. Robson
Hypericum mexicanum L.
Hypericum millefolium Urb. & Ekman
Hypericum myricariifolium Hieron.
Hypericum papillosum N.Robson
Hypericum parallelum N. Robson
Hypericum paramitanum N.Robson
Hypericum peninsulare Eastw.
Hypericum phellos Gleason
Hypericum pimeleoides Planch. & Linden ex Triana & Planch.
Hypericum piriai Arechav.
Hypericum prietoi N. Robson
Hypericum prostratum Cuatrec.
Hypericum pseudobrathys Turcz.
Hypericum pycnophyllum Urb.
Hypericum quitense R.Keller
Hypericum radicans N. Robson
Hypericum recurvum N. Robson
Hypericum roraimense Gleason
Hypericum rubritinctum N. Robson
Hypericum ruscoides Cuatrec.
Hypericum sabiniforme Trevis.
Hypericum selaginella N. Robson
Hypericum simonsii N.Robson
Hypericum sprucei N. Robson
Hypericum stenopetalum Turcz.
Hypericum strictum Kunth
Hypericum struthiolifolium Juss.
Hypericum stuebelii Hieron.
Hypericum styphelioides A.Rich.
Hypericum terrae-firmae Sprague & L.Riley
Hypericum tetrastichum Cuatrec.
Hypericum thuyoides Kunth
Hypericum valleanum N. Robson
Hypericum woodianum N. Robson
Hypericum wurdackii N. Robson

Bupleuroides

Bupleuroides Stef. contains only a single species that is a small perennial herb.

Hypericum bupleuroides Griseb.

Campylopus

Campylopus Boiss. contains only Hypericum cerastioides.

Hypericum cerastioides (Spach) N. Robson

Camplyosporus

Campylosporus (Spach) R. Keller

Hypericum balfourii N.Robson
Hypericum bequaertii De Wild.
Hypericum cerastioides (Spach) N.Robson
Hypericum gnidiifolium A.Rich.
Hypericum lanceolatum Lam.
Hypericum madagascariense (Spach) Steud.
Hypericum quartinianum A.Rich.
Hypericum revolutum Vahl
Hypericum roeperianum Schimp. ex A.Rich.
Hypericum socotranum R.D.Good
Hypericum synstylum N.Robson

Concinna

Hypericum concinnum Benth.

Coridium

Hypericum asperuloides Czern. ex Turcz.
Hypericum coris L.
Hypericum empetrifolium Willd.
Hypericum ericoides L.
Hypericum jovis Greuter

Crossophyllum

Hypericum adenotrichum Spach
Hypericum aucheri Jaub. & Spach
Hypericum orientale L.
Hypericum thasium Griseb.

Drosocarpium

Drosocarpium Spach contains small perennial herbs that are found near the Mediterranean.

Hypericum alpigenum Kit.
Hypericum ambiguum Elliott
Hypericum barbatum Jacq.
Hypericum bithynicum Boiss.
Hypericum confusum Rose
Hypericum montbretii Spach
Hypericum perfoliatum L.
Hypericum × reinosae A.Ramos
Hypericum richeri Vill.
Hypericum rumeliacum Boiss.
Hypericum spruneri Boiss.
Hypericum trichocaulon Boiss. & Heldr.
Hypericum umbellatum A.Kern.
Hypericum vesiculosum Griseb.

Elodeoida

Elodeoida N. Robson contains 1 m tall perennial herbs.

Hypericum austroyunnanicum L.H. Wu & D.P. Yang
Hypericum elodeoides Choisy
Hypericum hubeiense L.H. Wu & D.P. Yang
Hypericum kingdonii N.Robson
Hypericum petiolulatum Hook.f. & Thomson ex Dyer
Hypericum seniawinii Maxim.

Graveolentia

Graveolentia N. Robson is a diverse section whose type species is H. graveolens.

Hypericum collinum Schltdl. & Cham.
Hypericum epigeium R.Keller
Hypericum formosum Kunth
Hypericum graveolens Buckley
Hypericum macvaughii N.Robson
Hypericum × mitchellianum Rydb.
Hypericum oaxacanum R. Keller
Hypericum pringlei S.Watson
Hypericum pseudomaculatum Bush
Hypericum punctatum Lam.

Heterophylla

Heterophylla N. Robson contains a single shrublets measuring up to 25 cm tall.

Hypericum heterophyllum

Hirtella

Hirtella Stef. contains perennial herbs averaging 80 cm in height.

Hypericum amblysepalum Hochst.
Hypericum apiculatum (N.Robson) Sennikov
Hypericum apricum Kar. & Kir.
Hypericum asperulum Jaub. & Spach
Hypericum callithyrsum Coss.
Hypericum davisii N.Robson
Hypericum elongatum Ledeb. ex Rchb.
Hypericum hedgei N.Robson
Hypericum helianthemoides (Spach) Boiss.
Hypericum hirtellum (Spach) Boiss.
Hypericum karjaginii Rzazade
Hypericum libanoticum N.Robson
Hypericum lysimachioides Boiss. & Noë
Hypericum microcalycinum Boiss. & Heldr.
Hypericum olivieri (Spach) Boiss.
Hypericum pseudolaeve N.Robson
Hypericum retusum Aucher ex Jaub. & Spach
Hypericum salsolifolium Hand.-Mazz.
Hypericum scabroides N.Robson & Poulter
Hypericum scabrum L.
Hypericum sorgerae N.Robson
Hypericum spectabile Jaub. & Spach
Hypericum thymbrifolium Boiss. & Noë
Hypericum thymopsis Boiss.
Hypericum tymphresteum Boiss. & Spruner
Hypericum vermiculare Boiss. & Hausskn.

Humifusoideum

Humifusoideum R. Keller, also called Pulogensia, contains shrubs, subshrubs, and perennial herbs.

Hypericum bojerianum H. Perrier
Hypericum natalense J.M. Wood & M.S. Evans
Hypericum nokoense Ohwi
Hypericum peplidifolium A.Rich.
Hypericum wilmsii R. Keller

Hypericum

Hypericum, sometimes referred to as the "type section" of the genus, contains perennial herbs and very few suffrutices.

Hypericum asahinae Makino
Hypericum attenuatum Fisch. ex Choisy
Hypericum × desetangsii Lamotte
Hypericum elegans Stephan ex Willd.
Hypericum enshiense L.H. Wu & F.S. Wang
Hypericum erectum Thunb.
Hypericum faberi R.Keller
Hypericum furusei N.Robson
Hypericum gracillimum Koidz.
Hypericum hachijyoense Nakai
Hypericum hakonense Franch. & Sav.
Hypericum × hyugamontanum Kimura
Hypericum kamtschaticum Ledeb.
Hypericum kawaranum N.Robson
Hypericum kinashianum Koidz.
Hypericum kitamense (Y.Kimura) N.Robson
Hypericum kiusianum Koidz.
Hypericum kurodakeanum N.Robson
Hypericum × laschii A.Fröhl.
Hypericum maculatum Crantz
Hypericum × medium' Peterm.
Hypericum momoseanum Makino
Hypericum nakaii H.Koidz.
Hypericum nikkoense Makino
Hypericum nuporoense N.Robson
Hypericum oliganthum Franch. & Sav.
Hypericum ovalifolium Koidz.
Hypericum perforatum L.
Hypericum pibairense (Miyabe & Y.Kimura) N.Robson
Hypericum pseudoerectum N.Robson
Hypericum pseudopetiolatum R.Keller
Hypericum scouleri Hook.
Hypericum senanense Maxim.
Hypericum sikokumontanum Makino
Hypericum taihezanense Sasaki ex S.Suzuki
Hypericum tetrapterum Fr.
Hypericum triquetrifolium Turra
Hypericum undulatum Schousb. ex Willd.
Hypericum uniglandulosum Hausskn. ex Bornm.
Hypericum vulcanicum Koidz.
Hypericum watanabei N.Robson
Hypericum yamamotoanum H.Koidz.
Hypericum yamamotoi Miyabe & Kimura
Hypericum yezoense Maxim.

Inodora

Inodora Stef. contains one species, a shrub that grows from 1 to 5 meters tall.

Hypericum xylosteifolium (Spach) N.Robson

Monanthema

Monanthema N. Robson contains perennial herbs that grow up to 40 cm tall.

Hypericum daliense N.Robson
Hypericum himalaicum N.Robson
Hypericum ludlowii N.Robson
Hypericum monanthemum Hook.f. & Thomson ex Dyer
Hypericum subcordatum (R.Keller) N.Robson
Hypericum trigonum Hand.-Mazz.
Hypericum wightianum Wall. ex Wight & Arn.

Myrianda

Myriandra (Spach) R. Keller contains shrubs, shrublets, and perennial herbs that grow to be up to 4.5 m.

Hypericum adpressum W.P.C.Barton
Hypericum apocynifolium Small
Hypericum brachyphyllum (Spach) Steud.
Hypericum buckleyi M.A. Curtis
Hypericum chamaenerium (Spach) Steud.
Hypericum chapmanii W.P. Adams
Hypericum cistifolium Lam.
Hypericum crux-andreae (L.) Crantz
Hypericum densiflorum Pursh
Hypericum edisonianum (Small) W.P. Adams & N. Robson
Hypericum ellipticum Hook.
Hypericum exile W.P. Adams
Hypericum fasciculatum Lam.
Hypericum frondosum Michx.
Hypericum galioides Lam.
Hypericum glomeratum Small
Hypericum hypericoides (L.) Crantz
Hypericum kalmianum L.
Hypericum lissophloeus W.P. Adams
Hypericum lloydii (Svenson) P.B.Adams
Hypericum lobocarpum Gatt.
Hypericum longistylum Oliv.
Hypericum lydium Boiss.
Hypericum myrtifolium Lam.
Hypericum nitidum Lam.
Hypericum nudiflorum Michx. ex Willd.
Hypericum opacum Torr. & A. Gray
Hypericum prolificum L.
Hypericum suffruticosum W.P. Adams & N. Robson
Hypericum tetrapetalum Lam.

Oligostema

Oligostema (Boiss.) Stef. consista of perennial and annual herbs up to 75 cm tall.

Hypericum andjerinum Font Quer & Pau
Hypericum australe Ten.
Hypericum × caesariense Druce ex N.Robson
Hypericum humifusum L.
Hypericum kelleri Bald.
Hypericum linariifolium Vahl
Hypericum sphaerocarpum Michx.

Olympia

Olympia (Spach) Nymam contains four dwarf shrubs. They are found in Southeastern Europe and Turkey.

Hypericum auriculatum (N.Robson & Hub.-Mor.) N.Robson
Hypericum lycium (N.Robson & Hub.-Mor.) N.Robson
Hypericum olympicum L.
Hypericum polyphyllum Boiss. & Balansa

Origanifolia

Origanifolia Spach contains unique shrub-like perennial herbs.

Hypericum albiflorum (Hub.-Mor.) N.Robson
Hypericum aviculariifolium Jaub. & Spach
Hypericum bourgaei (Boiss.) N.Robson
Hypericum cymbiferum Boiss. & Balansa
Hypericum ichelense N.Robson
Hypericum imbricatum Poulter
Hypericum laxiflorum N.Robson
Hypericum leprosum Boiss.
Hypericum origanifolium Willd.
Hypericum papillare Boiss. & Heldr.
Hypericum salsugineum N.Robson & Hub.-Mor.
Hypericum trachyphyllum Griseb.
Hypericum uniflorum Boiss. & Heldr.

Psorophytum

Psorophytum (Spach) Nyman contains a single evergreen shrub. It grows from 1–5 meters tall.

Hypericum balearicum L.

Roscyna

Roscyna (Spach) R. Keller contains perennial herbs that grow up to 2 meters tall.

Hypericum ascyron L.
Hypericum przewalskii Maxim.

Sampsonia

Sampsonia N. Robson contains mainly perennial herbs that grow to be 80 cm.

Hypericum assamicum S.N.Biswas
Hypericum sampsonii Hance

Santomasia

Santomasia (N. Robson) N. Robson contains a single species.

Hypericum steyermarkii Standl.

Taeniocarpium

Taeniocarpium Jaub. & Spach contains small wiry perennial herbs up to 1.1 meters tall.

Hypericum armenum Jaub. & Spach
Hypericum crenulatum Boiss.
Hypericum fissurale Woronow
Hypericum havvae Güner
Hypericum hirsutum L.
Hypericum kotschyanum Boiss.
Hypericum linarioides Bosse
Hypericum malatyanum Peșmen
Hypericum marginatum Woronow
Hypericum neurocalycinum Boiss. & Heldr.
Hypericum nummularioides Trautv.
Hypericum nummularium L.
Hypericum peshmenii Yıld.
Hypericum pruinatum Boiss. & Balansa
Hypericum pseudorepens N.Robson
Hypericum pulchrum L.
Hypericum pumilio Bornm.
Hypericum saxifragum N.Robson & Hub.-Mor.
Hypericum taygeteum Quézel & Contandr.
Hypericum theodori Woronow
Hypericum thymifolium Banks & Sol.
Hypericum vaccinioides N.Robson
Hypericum venustum Fenzl

Takasagoya

Takasagoya (Y.Kimura) N. Robson contains deciduous shrubs and shrublets that grow up to 1.5 m tall.

Hypericum formosanum Maxim.
Hypericum geminiflorum Hemsl.
Hypericum nakamurai (Masam.) N.Robson
Hypericum senkakuinsulare Hatus.
Hypericum subalatum Hayata

Triadenoides

Triadenoides Jaub. & Spach contains shrubs and shrublets that can grow to be 0.8 m tall.

Hypericum fieriense N. Robson
Hypericum haplophylloides Halácsy & Bald.
Hypericum musadoganii Yıld.
Hypericum pallens Banks & Solander
Hypericum scopulorum Balf.f.
Hypericum ternatum Poulter
Hypericum tortuosum Balf.f.

Trigynobrathys

Trigynobrathys (Y. Kimura) N. Robson contains shrubs and subshrubs as well as annual and perennial herbs that are very diverse in size and shape.

Hypericum anagalloides Cham. & Schltdl.
Hypericum aphyllum Lundell
Hypericum arenarioides A. Rich.
Hypericum boreale (Britton) H. P. Bicknell
Hypericum brasiliense Choisy
Hypericum brevistylum Choisy
Hypericum caespitosum Cham. & Schltdl.
Hypericum campestre Cham. & Schltdl.
Hypericum canadense L.
Hypericum caprifoliatum Cham. & Schltdl.
Hypericum carinatum Griseb.
Hypericum cavernicola L.B. Sm.
Hypericum connatum Lam.
Hypericum cordatum (Vell.) N. Robson
Hypericum cumulicola (Small) P.B. Adams
Hypericum denticulatum Walter
Hypericum denudatum A. St.-Hil.
Hypericum × dissimulatum E.P. Bicknell
Hypericum dolabriforme Vent.
Hypericum erythreae (Spach) Steud.
Hypericum globuliferum R. Keller
Hypericum gramineum G.Forst.
Hypericum gymnanthum Engelm. & A.Gray
Hypericum harperi R.Keller
Hypericum humbertii Staner
Hypericum japonicum Thunb.
Hypericum killipii N. Robson
Hypericum lalandii Choisy
Hypericum legrandii L.B. Sm.
Hypericum linoides A. St.-Hil.
Hypericum lorentzianum Gilg ex R. Keller
Hypericum majus (A.Gray) Britton
Hypericum microlicioides L.B. Sm.
Hypericum microsepalum (Torr. & A. Gray) A. Gray ex S. Watson
Hypericum monadenum N.Robson
Hypericum moranense Kunth
Hypericum mutilum L.
Hypericum myrianthum Cham. & Schltdl.
Hypericum oligandrum Milne-Redh.
Hypericum parvulum Greene
Hypericum pauciflorum Kunth
Hypericum paucifolium S. Watson
Hypericum pedersenii N.Robson
Hypericum philonotis Schltdl. & Cham.
Hypericum pleiostylum C. Rodr. Jim.
Hypericum polyanthemum Klotzsch ex Reichardt
Hypericum pratense Schltdl. & Cham.
Hypericum pumilum Sessé & Moc.
Hypericum relictum N. Robson
Hypericum rigidum A. St.-Hil.
Hypericum salvadorense N. Robson
Hypericum scioanum Chiov.
Hypericum setosum L.
Hypericum silenoides Juss.
Hypericum teretiusculum A. St.-Hil.
Hypericum ternum A. St.-Hil.
Hypericum thesiifolium Kunth
Hypericum virgatum Lam.

Tripentas

Tripentas (Casp.) N. Robson contains long-stemmed perennial herbs.

Hypericum elodes L.

Umbraculoides

Umbraculoides N. Robson contains a single species by its own name.

Hypericum umbraculoides N.Robson

Webbia

Webbia (Spach) R. Keller contains a large deciduous shrub that can grow up to 4 meters tall.

Hypericum canariense L.

Unspecified

Hypericum callinum Rose
Hypericum × inodorum Mill.
Hypericum iwate-littorale H.Koidz.
Hypericum nagasawae Hayata
Hypericum rudiflorum Michx.
Hypericum tubulosum Walter

Bibliography

Studies in the genus Hypericum L. (Guttiferae)

References

↑ Le Roy Abrams & Roxana Stinchfield Ferris (1923). An Illustrated Flora of the Pacific States: Geraniaceae to Scrophulariaceae, geraniums to figworts. An Illustrated Flora of the Pacific States: Washington, Oregon, and California. 3. Stanford University Press. p. 116. ISBN 9780804700054.
↑ "Hypericum". The Plant List. Retrieved 19 February 2016.
↑ Hypericum MySpecies Retrieved March 4, 2016
↑ CABI.org Retrieved 9/7/16
↑ Robson, N.K.B. (1977) Studies in the genus Hypericum L. (Guttiferae) 1. Infrageneric classification Bulletin of the British Museum (Natural History), Botany 5: 293–355.
↑ Molecular phylogenetics and morphological evolution of St. John's wort (Hypericum; Hypericaceae) by Nürk NM, Madriñán S, Carine MA, Chase MW, Blattner FR

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