Temperate deciduous forest

Temperate forest in Germany just as the leaf canopy is opening

Temperate deciduous forests or temperate broad-leaf forests are dominated by trees that lose their leaves each year. They are found in areas with warm, moist summers and mild winters.[1] The three major areas of this forest type occur in the Northern Hemisphere: eastern North America, East Asia,[2] and Europe. Smaller areas occur in Australasia and southern South America.[3] Examples of typical trees in the Northern Hemisphere's deciduous forests include oak, maple, beech, and elm, while in Southern Hemisphere, trees of the genus Nothofagus dominate this type of forest. The diversity of tree species is higher in regions where the winter is milder, and also in mountainous regions that provide an array of soil types and microclimates.[4] One of the world's great protected examples of this forest type is found in Great Smoky Mountains National Park.[5][6]

Ecology

Bloodroot, like many other spring ephemerals, flowers in the spring before the forest canopy appears

The principal factor operating in these forests is the seasonal appearance and disappearance of the canopy.[7] Shade from the canopy limits the growth of many kinds of plants. Many species that are typical of these forests time their growth and flowering to the short period just before the canopy opens; hence they are known as spring ephemerals. Examples include trilliums and bloodroot. Most spring ephemerals are insect-pollinated, and the seeds themselves are often transported by ants,[8] a mode of dispersal known as myrmecochory. There are a smaller number of species able to grow under the canopy, and even a few that grow during the period when leaves are being lost. The average yearly precipitation is 30 - 60 inches (75 – 150 cm). Temperate deciduous forests have a great variety of plant species. Most have three levels of plants. Lichen, moss, ferns, wildflowers and other small plants can be found on the forest floor. Shrubs fill in the middle level and hardwood trees like maple, oak, birch, magnolia, sweet gum and beech make up the third level. Birds like broad-winged hawks, cardinals, snowy owls, and pileated woodpeckers are also found in this biome. Mammals include white-tailed deer, raccoons, opossums, porcupines and red foxes. Animals that live in the temperate deciduous forest must be able to adapt to the changing seasons.

Many understory plants have leaf adaptions to cope with low light levels, and the need to exploit moving flecks of light on the forest floor.[9] A few, such as Indian pipe and Corallorhiza orchids, have adapted to the shade by parasitism. The trees similarly are controlled by shade. Most tree seedlings require small gaps when trees fall and make space in order to regenerate. A few require larger gaps such as those produced by windstorms.[10] Gradients of soil moisture, soil depth, elevation and aspect control the distribution of many trees, shrubs and herbaceous species.[11] Some require unusual conditions such as steep slopes, infertile soil, and drought to escape competition from the more common tree species.[12]

Pileated woodpeckers depend upon dead or dying trees as a source of food and for constructing their nests.

Many migratory birds time their arrival to coincide with the opening of the canopy, which provides the insects that are their principal food sources for raising young. The spring warblers of North America are a typical example; see, for example, the black-throated blue warbler. Owing to the availability of wood from standing and fallen trees, woodpeckers are frequently found. The pileated woodpecker is a typical large species. Fallen wood, known as coarse woody debris, provides shelter for many kinds of amphibians, particularly salamanders. Many well-known animals live in this kind of forest; a few examples include squirrels, which are an important canopy species, and bears, which hibernate in the winter. The top predators in deciduous forest were once wolves and cougars, along with species of weasel like the fisher.

Human effects

Humans have often colonized areas in the temperate deciduous forest. They have harvested wood for timber and charcoal.[13] During the settlement of North America, potash made from tree ashes were exported back to Europe as fertilizer. As a result, less than one quarter of original forests remain. Many forests are now small fragments dissected by fields and roads; these islands of green often differ substantially from the original forests, particularly along the edges.[14][15] The introduction of exotic diseases continues to be a threat to forest trees, and hence, the forest;[16] examples include the loss of chestnut and elm. At the same time, species like deer, which are clearing rather than true forest animals, have expanded their range and proliferated in these altered landscapes.[17] Large deer populations have deleterious effects on tree regeneration overall, but particularly for edible species including yew, yellow birch and hemlock. Deer grazing also has significant negative effects on the number and kind of herbaceous flowering plants.[18] The continuing pressure to increase deer populations, and the continued killing of top carnivores, suggests that overgrazing by deer will continue to be a significant forest conservation problem. Objective criteria for the restoration of deciduous forest include large trees, coarse woody debris, spring ephemeral, and top predators.[19]

Gallery

See also

References

  1. ↑ Archibold, O. W. 1995. Ecology of World Vegetation, London: Chapman and Hall.
  2. ↑ Wen, J. 1999. Evolution of eastern Asian and eastern North American disjunct distributions in flowering plants. Annual Review of Ecology and Systematics 30:421-455
  3. ↑ Archibold, O. W. 1995. Ecology of World Vegetation. London: Chapman and Hall. Figure 6.1
  4. ↑ Keddy, P.A. 2007, Plants and Vegetation: Origins, Processes, Consequences, Cambridge University Press, Cambridge, UK.
  5. ↑ Braun, E. L. 1950, Deciduous Forests of Eastern North America, New York: Hafner
  6. ↑ Whittaker, R. H. 1956. Vegetation of the Great Smoky Mountains. Ecological Monographs 26: 1–79.
  7. ↑ Braun, E. L. 1950. Deciduous Forests of Eastern North America. New York: Hafner.
  8. ↑ Keddy, P.A. 2007. Plants and Vegetation: Origins, Processes, Consequences. Cambridge University Press, Cambridge, UK.
  9. ↑ Archibold, O. W. 1995. Ecology of World V1995. Physiological Plant Ecology: Ecophysiology and Stress Physiology of Functional Groups. 3rd edn. New York: Springer-Verlag.
  10. ↑ Keddy, P.A. 2007. Plants and Vegetation: Origins, Processes, Consequences. Cambridge University Press, Cambridge, UK.
  11. ↑ Whittaker, R. H. 1956. Vegetation of the Great Smoky Mountains. Ecological Monographs 26: 1–79.
  12. ↑ Keddy, P.A. and P. MacLellan. 1990. Centrifugal organization in forests. Oikos 58: 75-84.
  13. ↑ Hughes, J. D. 1982. Deforestation, erosion, and forest management in ancient Greece and Rome. Journal of Forest History 26: 60–75.
  14. ↑ Wilcove, D. S., C. H. McLellan, and A. P. Dobson. 1986. Habitat fragmentation in the temperate zone. pp. 237–256. In M. E. Soul´e (ed.) Conservation B; the Science of Scarcity and Diversity. Sunderland: Sinauer Associates.
  15. ↑ Harris, L. D. 1984. The Fragmented Forest: Island Biogeography Theory and the Preservation of Biotic Diversity. Chicago: University of Chicago Press.
  16. ↑ Little, C. E. 1995. The Dying of the Trees: The Pandemic in America’s Forests. New York: Penguin Books.
  17. ↑ Latham, R. E., J. Beyea, M. Benner, C.A. Dunn, M. A. Fajvan, R.R. Freed, M. Grund, S. B. Horsley, A. F. Rhoads, and B. P. Shissler. 2005. Managing White-tailed Deer in Forest Habitat from an Ecosystem Perspective: Pennsylvania Case Study. Harrisburg: Audubon Pennsylvania and Pennsylvania Habitat Alliance.
  18. ↑ Latham, R. E., J. Beyea, M. Benner, C.A. Dunn, M. A. Fajvan, R.R. Freed, M. Grund, S. B. Horsley, A. F. Rhoads, and B. P. Shissler. 2005. Managing White-tailed Deer in Forest Habitat from an Ecosystem Perspective: Pennsylvania Case Study. Harrisburg: Audubon Pennsylvania and Pennsylvania Habitat Alliance.
  19. ↑ Keddy, P.A. and C. G.Drummond. 1996. Ecological properties for the evaluation, management, and restoration of temperate deciduous forest ecosystems. Ecological Applications 6: 748–762.

External links

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