Kaatedocus

Kaatedocus Temporal range: Late Jurassic, 155 Ma PreЄ Є O S D C P T J K Pg N ↓
Mounted skeleton cast, Museum voor Natuurwetenschappen in Brussels, Belgium
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Clade:	Dinosauria
Order:	Saurischia
Suborder:	†Sauropodomorpha
Clade:	†Neosauropoda
Family:	†Diplodocidae
Subfamily:	†Diplodocinae
Genus:	†Kaatedocus Tschopp & Mateus, 2012
Type species
†Kaatedocus siberi Tschopp & Mateus, 2012

Kaatedocus is a genus of diplodocine flagellicaudatan sauropod known from the middle Late Jurassic (Kimmeridgian stage) of northern Wyoming, United States. It is known from well-preserved skull and cervical vertebrae which were collected in the lower part of the Morrison Formation. The type and only species is Kaatedocus siberi, described in 2012 by Emanuel Tschopp and Octávio Mateus.^[1]

History

Skull SMA 0004 from two angles

In 1934, a team of the American Museum of Natural History (AMNH) headed by Barnum Brown, financed by the Sinclair Oil Corporation, uncovered about three thousand sauropod bones on the land of rancher Barker Howe near Shell, in Big Horn County. Plans for further excavations in 1935 had to be cancelled after Howe, convinced by the large publicity surrounding the find that the remains were very valuable, demanded higher payment. The bones would not be described and most of them were lost in a fire at the AMNH during the 1940s; other were thrown away in the 1960s after having rotted because of being stowed in a chicken run at Shell. Only about 10% of the fossils survived, among them a skull. They were generally interpreted as belonging to Barosaurus. In 1989 the site was reopened by Hans-Jakob Siber, the founder of the Swiss Aathal Dinosaur Museum. His team immediately adjacent to the old How Quarry discovered another 450 bones that became part of the collection of the Swiss museum.

Size comparison with a human

The finds included an exceptionally complete neck. In Switzerland, this became the subject of several lines of scientific investigation. In 2005 Daniela Schwarz studied the pneumatisation of the vertebrae by tomography scanning them with neutrons and X-rays.^[2] In 2010 Andreas Christian used the well-preserved vertebrae to support his hypothesis that sauropod necks were held in a rather upright position,^[3] which was confirmed by Armin Schmitt studying the vestibular system of Kaatedocus. In 2012 Tschopp used a scan to create a replica of the neck by means of a 3D-printer.^[4]

During the intense study of the fossils it became clear that they did not represent Barosaurus but a species new to science. In 2012 this was named Kaatedocus siberi, by the Swiss palaeontologist Emanuel Tschopp, who as a boy had visited the excavations, and his Portuguese colleague Octávio Mateus. The generic name combines a reference to the related form Diplodocus with a Crow Indian diminutive suffix ~kaate. The specific name honours Siber.^[1]

Classification

Skeleton mounted in rearing posture, Sauriermuseum Aathal

Cladogram of the Diplodocidae after Tschopp, Mateus, and Benson (2015).^[5]

Diplodocidae	Amphicoelias altus  Apatosaurinae  Unnamed species Apatosaurus ajax Apatosaurus louisae Brontosaurus excelsus Brontosaurus yahnahpin Brontosaurus parvus  Diplodocinae  Unnamed species Tornieria africana Supersaurus lourinhanensis Supersaurus vivianae Leinkupal laticauda Galeamopus hayi Diplodocus carnegii Diplodocus hallorum Kaatedocus siberi Barosaurus lentus

References

1. Tschopp, E.; Mateus, O. V. (2012). "The skull and neck of a new flagellicaudatan sauropod from the Morrison Formation and its implication for the evolution and ontogeny of diplodocid dinosaurs". Journal of Systematic Palaeontology. 11: 1. doi:10.1080/14772019.2012.746589.
2. Schwarz, D.; Meyer, C. A.; Lehmann, E. H.; Vontobel, P.; Bongartz, G. (2005). "Testing Neutron tomography in comparison with X-ray computed tomography as a technique for the investigation of the internal structure of sauropod vertebrae and ribs". Palaeontologia Electronica. 8 (2): 1–11.
3. Christian, A. & Dzemski, G., 2011, "Neck posture in sauropods", pp: 251–260 in: Klein, N., Remes, K., Gee, C. G. & Sander, P. M. (eds.). Biology of the sauropod dinosaurs: Understanding the life of giants. Indiana University Press, Bloomington, 2011
4. Emanuel Tschopp & Gordon Dzemski, 2012, "3-Dimensional Reproduction Techniques to Preserve and Spread Paleontological Material – a Case Study with a Diplodocid Sauropod Neck", Journal of Paleontological Techniques 10
5. Tschopp, E.; Mateus, O. V.; Benson, R. B. J. (2015). "A specimen-level phylogenetic analysis and taxonomic revision of Diplodocidae (Dinosauria, Sauropoda)". PeerJ. 3: e857. doi:10.7717/peerj.857. PMC 4393826. PMID 25870766.