List of minor insects of Sri Lanka

Sri Lanka is a tropical island situated close to the southern tip of India. The invertebrate fauna is as large as it is common to other regions of the world. There are about 2 million species of arthropods found in the world, and still it is counting. So many new species are discover up to this time also. So it is very complicated and difficult to summarize the exact number of species found within a certain region.

The following list is about some minor insect orders recorded in Sri Lanka.

Webspinners

Phylum: Arthropoda
Class: Insecta
Order: Embioptera

The order Embioptera, commonly known as webspinners, are a small group of mostly tropical and subtropical insects, classified under the subclass Pterygota. The order has also been referred to as Embiodea or Embiidina.[1] The name Embioptera ("lively wings") comes from Greek, εμβιος, embios meaning "lively" and πτερον, pteron meaning "wing", a name that has not been considered to be particularly descriptive for this group of fliers,[2] perhaps instead referring to their remarkable speed of movement both forward and backward.[3]

Over 360 embiopteran species have been described,[2][4] along with estimates of around 2000 species being in existence today.[5] There is some debate as to the exact phylogenetic classification of Embioptera, with the order having been classed as a sister group to both orders Zoraptera,[2][6] and Phasmatodea,[7]

The following list provide the lacewings currently identified in Sri Lanka. Only four species found from this order within the country.[8][9][10][11][12]

Endemic species are denoted as E.

Family: Oligotomidae

Booklice

Phylum: Arthropoda
Class: Insecta
Order: Psocoptera

Psocoptera are an order of insects that are commonly known as booklice, barklice or barkflies.[17] They first appeared in the Permian period, 295–248 million years ago. They are often regarded as the most primitive of the hemipteroids.[18] Their name originates from the Greek word ψῶχος, psokos meaning gnawed or rubbed and πτερά, ptera meaning wings.[19] There are more than 5,500 species in 41 families in three suborders. Many of these species have only been described in recent years.[20]

The Order Psocoptera is divided into three suborders. According to checklists by Smithers in 1967 and New in 1977, there are 67 accepted species of booklice found from Sri Lanka.[8] The checklist of New was advanced with the full description of two subfamilies Epipsocidae and Pseudocaeciliidae, from Sri Lanka.[9][21][22]

Family: Amphientomidae - Tropical barklice

  • Tineomorpha greeniana
  • Paramphientomum nietneri
  • Seopsis metallops
  • Seopsis superba
  • Seopsis vasantasena
  • Syllysis caudata
  • Syllysis erato
  • Syllysis ritusamhara

Family: Amphipsocidae - Hairy-winged barklice

Family: Archipsocidae - Ancient barklice

Family: Caeciliusidae - Lizard barklice

  • Caecilius
  • Dypsocus coleoptratus
  • Isophanes palliatus
  • Coryphosmila dolabrata
  • Valenzuela aridus
  • Valenzuela maculistigma
  • Enderleinella ceylonica

Family: Calopsocidae - Common barklice

Family: Ectopsocidae - Outer barklice

Family: Elipsocidae - Damp barklice

Family: Epipsocidae - Elliptical barklice

Family: Hemipsocidae - Leaf litter barklice

Family: Lepidopsocidae - Scaly-winged barklice

  • Lepium luridum
  • Nepticulomima chalcomelas
  • Nepticulomima essigkeana
  • Nepticulomima sakuntala
  • Perientomum acutipenne
  • Perientomum argentatum
  • Perientomum ceylonicum
  • Perientomum chrysargyrium
  • Perientomum greeni
  • Perientomum gregarium
  • Perientomum morosum
  • Perientomum trichopteryx
  • Perientomum triste
  • Proentomum personatum
  • Soa flaviterminata
  • Echmepteryx mihira
  • Echmepteryx sericea
  • Scolopama halterata
  • Lepolepis ceylonica

Family: Liposcelididae - Booklice

Family: Myopsocidae - Mouse-like barklice

Family: Peripsocidae - Stout barklice

Family: Philotarsidae - Loving barklice

Family: Pseudocaeciliidae - False lizard barklice

  • Allopsocus medialis
  • Heterocaecilius ornatus
  • Mesocaecilius pictipennis
  • Ophiodopelma hieroglyphicum
  • Ophiodopelma multipunctatum
  • Pseudocaecilius cribrarius
  • Pseudocaecilius lanatus
  • Pseudocaecilius molestus
  • Pseudocaecilius ornatus
  • Pseudocaecilius paraornatus
  • Pseudocaecilius zonatus

Family: Psocidae - Common barklice

Family: Stenopsocidae - Narrow barklice

Thrips

Phylum: Arthropoda
Class: Insecta
Order: Thysanoptera

Thrips (order Thysanoptera) are minute, slender insects with fringed wings (thus the scientific name. Other common names for thrips include thunderflies, thunderbugs, storm flies, thunderblights, storm bugs, corn flies and corn lice. Thrips species feed on a large variety of plants and animals by puncturing them and sucking up the contents. A large number of thrips species are considered pests, because they feed on plants with commercial value. Some species of thrips feed on other insects or mites and are considered beneficial, while some feed on fungal spores or pollen. Approximately 6,000 species have been described. Thrips are generally tiny (1 mm long or less) and are not good flyers , although they can be carried long distances by the wind. In the right conditions, like indoor grow rooms or greenhouses, many species can exponentially increase in population size and form large swarms because of a lack of natural predators, making them an irritation to humans.

The first comprehensive detailed work on Sri Lankan thrip fauna came through Schmutz in 1913. His checklist stood for more than 70 years with 43 new species.[8] In 1997, Oda et al. rediscovered and updated the thrip diversity, but with small collections from Sri Lanka.[9] The most recent work was done by Wijerathna, and he listed 16 species of thrips from 28 crops across the island. Currently, thrips documented within Sri Lanka included to 3 families - Aeolothripidae, Thripidae, and Phlaeothripidae, with 46 genera and 78 species.[23][24]

Family: Aeolothripidae - Predatory thrips

Family: Thripidae - Common thrips

  • Caliothrips graminicola
  • Caliothrips indicus
  • Copidothrips octarticulatus
  • Helionothrips brunneipennis
  • Heliothrips haemorrhoidalis
  • Noathrips prakashi
  • Panchaetothrips indicus
  • Parthenothrips dracaenae
  • Phibalothrips peringueyi
  • Retithrips syriacus
  • Rhipiphorothrips cruentatus
  • Rhipiphorothrips pulchellus
  • Selenothrips rubrocinctus
  • Tryphactothrips rutherfordi
  • Pseudodendrothrips ornatissimus
  • Dendrothrips sexmaculatus
  • Anaphothrips sudanensis
  • Bolacothrips striatopennatus
  • Bregmatothrips brachycephalus
  • Chaetanaphothrips signipennis
  • Deuterobrachythrips lineatus
  • Frankliniella occidentalis
  • Frankliniella schultzei
  • Megalurothrips distalis
  • Megalurothrips typicus
  • Megalurothrips usitatus
  • Microcephalothrips abdominalis
  • Neohydatothrips samayunkur
  • Sciothrips cardamomi
  • Scirtothrips dorsalis
  • Stenchaetothrips biformis
  • Thrips coloratus
  • Thrips flavus
  • Thrips Jlorum
  • Thrips hawaiiensis
  • Thrips longalatus
  • Thrips palmi
  • Thrips simplex
  • Thrips tabaci

Family: Phlaeothripidae - Tube-tailed thrips

  • Bactrothrips idolomorphus
  • Diaphorothrips unguipes
  • Dinothrips spinosus
  • Dinothrips sumatrensi
  • Elaphrothrips denticollis
  • Elaphrothrips greeni
  • Elaphrothrips malayensis
  • Elaphrothrips procer
  • Ethirothrips angusticornis
  • Ethirothrips indicus
  • Ethirothrips obscurus
  • Ethirothrips stenomelas
  • Ethirothrips watsoni
  • Ischyrothrips crassus
  • Mecynothrips simplex
  • Neosomerinthothrips affinis
  • Neosomerinthothrips fructuum
  • Aleurodothrips fasciapennis
  • Androthrips flavipes
  • Chromatothrips annulicornis
  • Chromatothrips fasciatus
  • Chromatothrips plantaginis
  • Ecacanthothrips tibialis
  • Eumorphothrips albicornis
  • Gigantothrips schenklingi
  • Gigantothrips tibialis
  • Haplothrips ceylonicus
  • Haplothrips ganglbaueri
  • Haplothrips gowdeyi
  • Haplothrips terminalis
  • Liothrips floridensis
  • Liothrips karnyi
  • Liothrips mirabilis
  • Liothrips tropicus
  • Liothrips vaneeckei
  • Teuchothrips brevis
  • Teuchothrips longus
  • Trichinothrips breviceps

Fleas

Phylum: Arthropoda
Class: Insecta
Order: Siphonaptera

Fleas are insects that form the order Siphonaptera. They are wingless, with mouthparts adapted for piercing skin and sucking blood. Fleas are external parasites, living by hematophagy off the blood of mammals and birds. Over 2,000 species have been described worldwide.[25]

The following list provide the fleas found in Sri Lanka.[9] The first checklist of fleas in Sri Lanka was done by Iyengar in 1973.[8] 20 species are recognized, more taxonomic study is required.[26] The fleas studies were almost confined to parasitic sections, where W. W. A. Phillips documented 11 species of fleas in 1980.

Family: Ceratophyllidae

Family: Ischnopsyllidae - Bat fleas

Family: Pulicidae - Cat fleas

Family: Stivaliidae

Caddisflies

Phylum: Arthropoda
Class: Insecta
Order: Trichoptera

The caddisflies are an order, Trichoptera, of insects with approximately 7,000 described species.[27] Also called sedge-flies or rail-flies, they are small moth-like insects having two pairs of hairy membranous wings. They are closely related to Lepidoptera (moths and butterflies) which have scales on their wings, and the two orders together form the superorder Amphiesmenoptera. Caddisflies have aquatic larvae and are found in a wide variety of habitats such as streams, rivers, lakes, ponds, spring seeps, and temporary waters (vernal pools).[28] The larvae of many species use silk to make protective cases of gravel, sand, twigs or other debris.

The caddisfly diversity in Sri Lanka is fairly studied from British times to present day. However, the first comprehensive work was done by Schmid in 1958.[29] Then in 1973, Malicky updated the checklist.[9] Currently 188 number of caddisfly species belongs to 18 families are identified from Sri Lanka.[8][30][31][32][33]

Family: Anomalopsychidae

Family: Calamoceratidae

Family: Dipseudopsidae

Family: Ecnomidae

  • Ecnomus ceylanicus
  • Ecnomus chusie
  • Ecnomus dutthangamani
  • Ecnomus helakanda
  • Ecnomus hinayana
  • Ecnomus indicus
  • Ecnomus lohaprasada
  • Ecnomus saddhatissa
  • Ecnomus tenellus
  • Ecnomus vaharika
  • Ecnomus vahasaba

Family: Goeridae

Family: Glossosomatidae - Little black caddisflies

  • Agapetus anuragoda
  • Agapetus ayodhia
  • Agapetus hanumata
  • Agapetus kithmalie
  • Agapetus kumudumalie
  • Agapetus rama
  • Agapetus rawana
  • Agapetus rudis
  • Agapetus sita

Family: Helicopsychidae - Snail-case caddisflies

Family: Hydrobiosidae

Family: Hydropsychidae - Net-spinning caddisflies

  • Amphipsyche meridiana
  • Amphipsyche sinhala
  • Cheumatopsyche curvata
  • Cheumatopsyche galahittigama
  • Cheumatopsyche galapitikanda
  • Cheumatopsyche kirimaduwa
  • Macrostactobia elawalikanda
  • Macrostemum indistinctum
  • Macrostemum kolenati
  • Macrostemum multifarium
  • Macrostemum nebulosum
  • Macrostemum pseudoneura
  • Macrostemum splendidum
  • Molanna taprobane
  • Oestropsyche vitrina
  • Potamyia nikalandugola
  • Pseudoleptonema godapitigama
  • Pseudoleptonema kalukandama

Family: Hydroptilidae - Micro caddisflies

  • Chrysotrichia aranuwa
  • Chrysotrichia dotalugola
  • Chrysotrichia hapitigola
  • Chrysotrichia hatnagola
  • Chrysotrichia porsawan
  • Chrysotrichia siriya
  • Hydroptila dikirilagoda
  • Hydroptila furcata
  • Hydroptila hemeli
  • Hydroptila kirilawela
  • Hydroptila kurukepitiya
  • Hydroptila mitirigalla
  • Hydroptila sumanmalie
  • Hydroptila upulmalie
  • Nietnerella hageni
  • Nyctiophylax abaya
  • Nyctiophylax devanampriya
  • Nyctiophylax hettigegama
  • Nyctiophylax tallawakanda
  • Nyctiophylax vetulya
  • Orthotrichia guruluhela
  • Orthotrichia hinipitigola
  • Orthotrichia indica
  • Orthotrichia litoralis
  • Orthotrichia udawarama
  • Oxyethira bogambara
  • Oxyethira galekoluma
  • Oxyethira incana
  • Oxyethira rachanee
  • Paduniella ceylanica
  • Paduniella mahanawana
  • Paduniella mahindra
  • Paduniella methinee
  • Paduniella pandya
  • Paduniella sanghamittra
  • Paduniella siveci
  • Paduniella subhakara
  • Paduniella thitima
  • Paduniella vattagamani
  • Paduniella vikramasinha
  • Parastactobia talakalahena
  • Plethus amogawarsa
  • Plethus bodikatuwa
  • Plethus cilamegha
  • Plethus cursitans
  • Plethus udawasadenna
  • Plethus vajrabodhi
  • Rhyacophila castanea
  • Stactobia fischeri

Family: Lepidostomatidae - Bizarre caddisflies

Family: Leptoceridae - Long-horned caddisflies

  • Adicella agastya
  • Adicella biramosa
  • Ceraclea isurumuniya
  • Gunungiella madakumbura
  • Gunungiella nimitra
  • Leptocerus anuradha
  • Leptocerus argentoniger
  • Leptocerus charopantaja
  • Leptocerus mahasena
  • Leptocerus mahawansa
  • Leptocerus parakum
  • Leptocerus posticus
  • Oecetis belihuloya
  • Oecetis biramosa
  • Oecetis ceylanica
  • Oecetis dhatusena
  • Oecetis fahieni
  • Oecetis hamata
  • Oecetis jacobsoni
  • Oecetis lingua
  • Oecetis malighawa
  • Oecetis meghadouta
  • Oecetis naravitta
  • Oecetis nerviciliata
  • Oecetis punctatissima
  • Oecetis sumanasara
  • Parasetodes respersellus
  • Setodes argentoaureus
  • Tagalopsyche brunnea
  • Triaenodes lankarama
  • Triaenodes ornatus
  • leiochiton suwannee
  • Trichosetodes argentolineatus
  • Trichosetodes meghawanabaya
  • Triplectides ceylanicus

Family: Limnephilidae - Northern caddisflies

Family: Odontoceridae - Mortarjoint casemakers

Family: Philopotamidae - Fingernet caddisflies

  • Chimarra actinifera
  • Chimarra akarawitta
  • Chimarra auriceps
  • Chimarra auricoma
  • Chimarra ceylanica
  • Chimarra circularis
  • Chimarra confusa
  • Chimarra godagama
  • Chimarra jiraprapa
  • Chimarra lankana
  • Chimarra lewisi
  • Chimarra mitis
  • Chimarra prisna
  • Chimarra sandhamma
  • Chimarra sepulchralis
  • Chimarra telihigola
  • Chimarra uvana
  • Chimarra wiharawela

Family: Polycentropodidae - Tube-maker caddisflies

  • Pahamunaya layagammeda
  • Polyplectropus amarawathi
  • Polyplectropus matadapaya
  • Polyplectropus nubigenus
  • Polyplectropus parakrama
  • Pseudoneureclipsis hataya
  • Pseudoneureclipsis maliboda
  • Pseudoneureclipsis narita
  • Pseudoneureclipsis nissanka
  • Pseudoneureclipsis thuparama
  • Pseudoneureclipsis watagoda
  • Pseudoneureclipsis wilaiwan
  • Pseudoneureclipsis yuwadee

Family: Psychomyiidae - Net-tube caddisflies

Family: Xiphocentronidae

Twisted-winged parasites

Phylum: Arthropoda
Class: Insecta
Order: Strepsiptera

Order Strepsiptera, commonly called, twisted-wing parasites, are an endopterygote order of insects. The order consists with nine extant families with about 600 species. Adults in most of their lives are spent as endoparasites in other insects, such as bees, wasps, leafhoppers, silverfish, and cockroaches.[34] Males have well-developed pair of hind-wings and reduced fore-wings. Females wingless and usually do not leave their hosts.[8]

The first scientific observation and detailed work on strepsipterans of Sri Lanka was done by Kathirithamby in 1994.[35][36] In 1997, Kifune discovered 20 strepsipterans from Sri Lanka, with 7 new species. All these new species genera are endemic to the country.[37][9][38][39]

Family: Corioxenidae

Family: Elenchidae

Family: Halictophagidae

Family: Mengenillidae

Family: Myrmecolacidae

Family: Stylopidae

References

  1. ↑ Donald J. Borror; Charles A. Triplehorn; Norman F. Johnson (1989). An Introduction to the Study of Insects (6th ed.). Harcourt Brace College Publishers. p. 247.
  2. 1 2 3 Michael S. Engel & David Grimaldi (2006). "The earliest webspinners (Insecta: Embiodea)" (PDF). American Museum Novitates. 3514: 1–22. doi:10.1206/0003-0082(2006)3514[1:tewie]2.0.co;2.
  3. ↑ Daniel Rains Wallace (2009). "Biologist Janice Edgerly-Rooks & the Extraordinary Embiids, Silken Choreographies". Santa Clara Magazine. Spring.
  4. ↑ Claudia Szumik (2008). "Phylogeny of embiopterans (Insecta)". Cladistics. 24: 993–1005. doi:10.1111/j.1096-0031.2008.00228.x.
  5. ↑ E. S. Ross (2000). "Contributions to the biosystematics of the insect order Embiidina. Part 1. Origin, relationships and integumental anatomy of the insect order Embiidina". Occasional Papers of the California Academy of Sciences. 149: 1–53.
  6. ↑ K. Yoshizawa (2007). "The Zoraptera problem: evidence for Zoraptera plus Embiodea from the wing base". Systematic Entomology. 32 (2): 197–204. doi:10.1111/j.1365-3113.2007.00379.x.
  7. ↑ Matthew D. Terry; Michael F. Whiting (2005). "Mantophasmatodea and phylogeny of the lower neopterous insects". Cladistics. 21 (3): 247–257. doi:10.1111/j.1096-0031.2005.00062.x.
  8. 1 2 3 4 5 6 Wijesekara, Anura and Wijesinghe, D.P. "HISTORY OF INSECT COLLECTION AND A REVIEW OF INSECT DIVERSITY IN SRI LANKA". Ceylon Journal of Science. p. 59. Retrieved 6 June 2016.
  9. 1 2 3 4 5 6 Bambaradeniya, Channa N. B. "The Fauna of Sri Lanka: Status of Taxonomy, Research, and Conservation". Amazon.com. Google books. Retrieved 23 January 2016.
  10. ↑ Chandra, Kailash. Dawn, Prosenjit. "First records of four species of webspinners (Insecta: Embioptera) from Chhattisgarh, India". Retrieved 8 June 2016.
  11. ↑ Ross, Edwards. "Embioptera from Sri Lanka (Ceylon)" (PDF). Retrieved 8 June 2016.
  12. ↑ Poolprasert, Pisit. "The Embiopteran Genus Oligotoma Westwood, 1837 (Embioptera: Oligotomidae), with Three New Recorded Species from Thailand" (PDF). Kasetsart J. (Nat. Sci.). Retrieved 8 June 2016.
  13. ↑ "species Oligotoma humbertiana (Saussure, 1896)". Embioptera Species File (Version 5.0/5.0). Retrieved 8 June 2016.
  14. ↑ "species Oligotoma saundersii (Westwood, 1837)". Embioptera Species File (Version 5.0/5.0). Retrieved 8 June 2016.
  15. ↑ "species Aposthonia ceylonica (Enderlein, 1912)". Embioptera Species File (Version 5.0/5.0). Retrieved 8 June 2016.
  16. ↑ "species Aposthonia minuscula (Enderlein, 1912)". Embioptera Species File (Version 5.0/5.0). Retrieved 8 June 2016.
  17. ↑ http://www.brc.ac.uk/schemes/barkfly/introduction.htm
  18. ↑ Christopher O'Toole (2002). Firefly Encyclopedia of Insects and Spiders. Toronto: Firefly Books. ISBN 1-55297-612-2.
  19. ↑ John R. Meyer (2005-03-05). "Psocoptera". North Carolina State University.
  20. ↑ Alfonso N. García Aldrete (2006). "New genera of Psocoptera (Insecta), from Mexico, Belize and Ecuador (Psoquillidae, Ptiloneuridae, Lachesillidae)" (PDF). Zootaxa. 1319: 1–14.
  21. ↑ New, T. R. "Epipsocidae and Pseudocaeciliidae (Psocoptera) from Sri Lanka". Oriental Insects Volume 11, Issue 3, 1977. Retrieved 8 June 2016.
  22. ↑ "Checklist of Psocoptera according to the World Catalogue (Lienhard & Smithers, 2002)". Natural History Museum of the City of Geneva. Department of Arthropodology and Entomology I. Retrieved 8 June 2016.
  23. ↑ Tillekaratne, Kalpana , Mound, Laurence A. , Zur Strassen, R. , Edirisinghe, Jayanthi P. "List of Thrips (Thysanoptera) recorded from Sri Lanka" (PDF). J. NatnSci.Foundation Sri Lanka. Retrieved 8 June 2016.
  24. ↑ Tillekaratne, Kalpana , Edirisinghe, J.P. , Gunatilleke, C.V.S. , Karunaratne, W. A. I. P. "Survey of thrips in Sri Lanka: A checklist of thrips species, their distribution and host plants" (PDF). University of Peradeniya. Retrieved 8 June 2016.
  25. ↑ Fleas: What They Are, What To Do D. L. Richman and P. G. Koehler, University of Florida IFAS Extension. Accessed 10 December 2010
  26. ↑ "Checklist of Siphanaptera in Sri Lanka". insectoid.info. Retrieved 8 June 2016.
  27. ↑ Keith Arthur John Wise (2016-02-22). "Caddisfly". Encyclopaedia Britannica.
  28. ↑ Glenn B. Wiggins, Larvae of the North American Caddisfly General (Trichoptera), 2nd. ed. (Toronto: University Press, 1996), p. 3
  29. ↑ "Insecta-Trichoptera checkList" (PDF). biodiversity.be. Retrieved 8 June 2016.
  30. ↑ "Diplectroninae Of Sri Lanka (Trichoptera : Hydropsychidae)". biodiversitylibrary.org. Retrieved 8 June 2016.
  31. ↑ "A new species of Anisocentropus (Trichoptera: Calamoceratidae) from Sri Lanka". Biodiversity library. Retrieved 8 June 2016.
  32. ↑ Chantaramongkol, P. "The Caddisflies of Sri Lanka". Proceedings of the Fifth International Symposium on Trichoptera Volume 39 of the series Series Entomologica. pp. 109–110. Retrieved 8 June 2016.
  33. ↑ Malicky, Hans. "Caddisflies (Trichoptera) from Parakrama Samudra, an ancient man-made lake in Sri Lanka". Limnology of Parakrama Samudra — Sri Lanka. Volume 12 of the series Developments in Hydrobiology. pp. 227–228. Retrieved 8 June 2016.
  34. ↑ Whiting, M. F (2003). "Strepsiptera". In Resh, V. H. & R. T. Cardé. Encyclopedia of Insects. Academic Press. pp. 1094–1096.
  35. ↑ Kathirithamby, Jeyaraney. "DESCRIPTIONS OF STREPSIPTERA (INSECTA) FROM SOUTHEAST ASIA, WITH A CHECKLIST OF THE GENERA AND SPECIES OCCURRING IN THE REGION" (PDF). Raffles bulletin of Zoology. Retrieved 9 June 2016.
  36. ↑ Kathirithamby, Jeyaraney. "Partial List of Strepsiptera Species". Tree of Life Project. Retrieved 9 June 2016.
  37. ↑ Kifune, Teiji. "RECORDS OF THE STREPSIPTERA OF SRI LANKA IN THE COLLECTION OF THE SMITHSONIAN INSTITUTION, WITH DESCRIPTIONS OF SEVEN NEW SPECIES (Notulae Strepsipterologicae-VI)" (PDF). ESAKIA. pp. 143–159. Retrieved 9 June 2016.
  38. ↑ "Zeylanica/Journal of South Asian Natural History > Volume 1 Number 1". wht.lk. Retrieved 9 June 2016.
  39. ↑ "THREE NEW SPECIES OF THE GENUS PARAXENOS (STREPSIPTERA : STYLOPIDAE) PARASITIC ON BEMBIX (HYMENOPTERA : SPHECIDAE) OF SRI LANKA AND AUSTRALIA IN THE COLLECTION OF THE SMITHSONIAN INSTITUTION (Notulae Strepsipterologicae-XVII)". ResearchGate. Retrieved 9 June 2016.
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