Bats described in the 2000s

This page is a list of species of the order Chiroptera discovered in the 2000s. See also parent page Mammals described in the 2000s.

2008

Desmalopex microleucopterus (2008)

A new species of flying fox found on Mindoro Island, Philippines.[1]

Triaenops pauliani (2008)

A new species discovered from the Picard Island (Aldabra Atoll, Seychelles).[2]

2007

Barbastella beijingensis (2007)

A new species found only in Fangshan District, about 100 km southwest of Beijing (China).[3]

Styloctenium mindorensis (2007)

This species of stripe-faced fruit bat is known only from the type locality in western Mindoro Island, Philippines.[4]

Hipposideros boeadii (2007)

A new species of Hipposideros described from South-East Sulawesi, Indonesia. It is currently only known from Rawa Aopa Watumohai National Park, an area of semi-disturbed lowland rainforest.[5]

Kerivoula krauensis (2007)

A new species of small Kerivoula described from peninsular Malaysia.[6]

2006

Eptesicus taddeii (2006)

A new species of Eptesicus from Atlantic Forest, Brazil. Zootaxa 2006[7]

Lonchophylla pattoni and L. cadenai (2006)

Two new species described from western South America. L. pattoni is known only from the type locality in the Amazon lowlands of southeastern Peru. L. cadenai was found only on the central Pacific coastal plain, Valle del Cauca, southwestern Colombia.[8]

Khasian Leaf-nosed Bat Hipposideros khasiana (2006)

A species of horseshoe bat from north-east India (Meghalaya state), proposed as distinct from the Intermediate Roundleaf Bat (H. larvatus) in 2006. It differs from H. larvatus in having a call at a lower frequency, and having longer ears and forearms.[9]

Phou Khao Khouay Leaf-nosed Bat Hipposideros khaokhouayensis (2006)

This is a species from Laos of the H. bicolor subgroup within the family Hipposideridae, with long brown dorsal fur and very light brownish ventral fur, large ears, relatively small and narrow nose-leaf without supplementary leaflets. It is further distinguished by cranial characters and the frequency of its echolocation calls.[10]

Marovaza House Bat Scotophilus marovaza (2006)

A new species of house bat discovered in 2004 and named in 2006. It was discovered living in palm-thatched roofs in the Madagascar village of Marovaza.[11]

Harpiola isodon (2006)

A new species was found in mountainous coniferous plantations and mixed forests of coniferous and broadleaf trees in Taiwan.[12]

Plecotus strelkovi (2006)

This new species inhabits the xeric mountain ranges in Central Asia (Afghanistan, China, Iran, Kazakhstan, Kyrgyzstan, Mongolia, Tajikistan).[13]

Carollia benkeithi (2006)

A species described from the lowland forests of eastern Peru and northwestern Bolivia.[14]

2005

Anoura fistulata (2005)

This new species inhabits montane cloud forests in the Andes of northern Ecuador, and the slopes of the Cordillera de Cóndor and Cordillera del Cutucú in southern Ecuador at elevations 1,300–1,890 m on the eastern and 2,000–2,275 m on the western slopes.[15]

Xeronycteris vieirai (2005)

A new genus and species in the tribe Lonchophyllini, subfamily Glossophaginae (Chiroptera: Phyllostomidae) was described based on the analysis of 4 specimens collected in 3 different localities in a semiarid area of northeastern Brazil.[16]

Greater monkey-faced bat Pteralopex flanneryi (2005)

This is the largest known monkey-faced bat, found on the northwestern Solomon Islands of Bougainville, Buka, Choiseul and Isabel, and some adjacent smaller islands. It has a forearm length of 159mm or more, and a condylobasal skull length of 71 mm or more. Its fur is black with occasional light tipping on the breast.[17]

Rhinolophus chiewkweeae (2005)

A new species classified in the R. pearsoni species group inhabits a dipterocarp forest of peninsular Malaysia.[18]

Lonchophylla orcesi (2005)

A species described from the Chocó region of Ecuador.[19]

Kock's Mouse-eared Bat Myotis dieteri (2005)

A very small Subsaharan Myotis, with a forearm length of 37 mm, brown dorsal and greyish ventral pelage.[20]

Harrison's Murine Bat Murina harrisoni (2005)

This bat was collected in a degraded gallery forest in Kirirom National Park in Cambodia. It differs from all other Murina species by its second upper incisor being shorter than the first, and from all but one by the insertion of its tail membrane on the base of the first toe.[21]

Scotophilus tandrefana (2005)

This new species is known from three localities in western Madagascar (Bemaraha, Mahabo, and Sarodrano).[22]

2004

Ansell's Epauletted Fruit Bat Epomophorus anselli (2004)

This new fruit bat was discovered in two bat collections from Malawi. The species is intermediate in size between the sympatric species E. labiatus and E. crypturus. Its wings are relatively broad, its tail membrane is narrow. In females, the 5th palatal ridge is partly between the first upper molars.[23]

Red or Chocolate Free-tailed Bat Chaerephon jobimena (2004)

This species was caught in northern Madagascar and consists of two typical colour morphs, a reddish one and a medium chocolate brown one.[24]

Kachin Woolly Bat Kerivoula kachinensis (2004)

This bat was found in forests in North Myanmar. It is characterized by the combination of its large size – it is larger than most other Asiatic Kerivoula species – and its unique flattened skull.[25]

Hanaki's Dwarf Bat Pipistrellus hanaki (2004)

This new dwarf bat from Libya differs from its nearest relatives P. pipistrellus and P. pygmaeus chromosomally, and by its larger skull and teeth, and a number of other characters.[26]

Lonchophylla chocoana (2004)

A new species currently known only from northwestern Ecuador and southwestern Colombia.[27]

Lophostoma yasuni (2004)

A new species is known only from the type locality in Yasuni National Park, Ecuador.[28]

Lophostoma aequatorialis (2004)

A new species from the Pacific coast of northwestern Ecuador (Esmeraldas, Los Ríos, and Pichincha provinces).[29]

Carollia manu (2004)

A new species is known from southeastern Peru and northern Bolivia.[30]

2003

Linduan Rousette Rousettus linduensis (2003)

This species was discovered in Lore Lindu National Park in Central Sulawesi, Indonesia.[31]

Shield-nosed Bat Hipposideros scutinares (2003)

This is a large leaf-nosed bat from Laos and Vietnam, with a known forearm length of 77.9–82.7 mm and a condylobasal length of 26.5–27.9 mm. It is intermediate between its two nearest relatives and further differs from these in a number of body and skull characters.[32]

2002

New Caledonian Long-eared Bat Nyctophilus nebulosus (2002)

This species is only known from the area of Nouméa on New Caledonia and is considered vulnerable by its author.[33]

Sowell's Short-tailed Bat Carollia sowelli (2002)

This species occurs from San Luis Potosi in Mexico to West Panama.[34]

Matses Big-eared Bat Micronycteris matses (2002)

Known from a single locality in Peru.[35]

Sakeji Horseshoe Bat Rhinolophus sakejiensis (2002)

A species discovered in Zambia.[36]

New Georgian Monkey-faced Bat Pteralopex taki (2002)

A species from New Georgia Island and Vangunu Island, Solomon Islands, considered by its author to be critically endangered.[37]

Sardinian Long-eared Bat Plecotus sardus (2002)

This bat is distinguished by its large tragus, its brownish dorsal pelage, its large thumb and thumb claw, and the form of its penis and penis bone.[38]

Rhinolophus ziama (2002)

A member of the R. maclaudi species group. Was described in 2002. This horseshoe bat species is known from a few specimens taken from Ziama Classified Forest in southeastern Guinea. It is also known from Liberia.[39]

Moa Island Fruit Bat Pteropus banakrisi (2002)

Was described as the smallest of Australian Pteropus species, but in 2004 Helgen found that the specimens on which P. banakrisi is based are all subadult individuals of the Black Flying Fox (P. alecto). The name banakrisi becomes a synonym.[40][41]

2001

Annami Myotis Myotis annamiticus (2001)

Only known from a single locality in Vietnam.[42]

Alcathoe Myotis Myotis alcathoe (2001)

This species is known from Greece, Hungary and France.[43]

Colombian Short-tailed Bat Carollia colombiana (2001)

This species is known only from the central Cordillera in Colombia.[44]

Antioquian Sac-winged Bat Saccopteryx antioquensis (2001)

A species discovered in the Central Cordillera of North Colombia.[45]

Nijhoff's Tube-nosed Bat Paranyctimene tenax (2001)

This fruit bat species was described in 2001, based on 1 specimen from Papua New Guinea, 7 from Vogelkop, Indonesia, and 1 from Waigeo, Indonesia. It is characterized by its long upper and lower canines and distinguished from its close and sympatric relative P. raptor by its larger size and stronger built.[46]

Curry's Butterfly Bat Glauconycteris curryae (2001)

This new butterfly bat from Cameroon and the Democratic Republic of Congo is distinct from other African species of its genus by the combination of the characters size, skull shape, fur colour, and the absence of spotting or reticulation. The authors named the bat G. curryi but later in 2001 the name was corrected to G. curryae.[47][48]

2000

Bale Long-eared Bat Plecotus balensis (2000)

This species was discovered in the Bale Mountains National Park of Ethiopia, at an altitude of 2760 m.[49]

Mistratoan Yellow-shouldered Bat Sturnira mistratensis (2000)

A species discovered in the western Andes of Colombia.[50]

Maendeleo Horseshoe Bat Rhinolophus maendeleo (2000)

Described on the basis of two specimens from Tanzania.[51]

References

  1. ↑ Esselstyn, J.A.; Garcia, H.J.D.; Saulog, M.G.; Heaney, L.R. (2008). "A New Species of Desmalopex (Pteropodidae) from the Philippines, with a Phylogenetic Analysis of the Pteropodini" (PDF). Journal of Mammalogy. 89 (4): 815–825. doi:10.1644/07-mamm-a-285.1.
  2. ↑ Goodman, S.M.; Ranivo, J. (2008). "A new species of Triaenops (Mammalia, Chiroptera, Hipposideridae) from Aldabra Atoll, Picard Island (Seychelles)" (PDF). Zoosystema. 30 (3): 681–693.
  3. ↑ Zhang, J.-S.; Han, N.-J.; Jones, G.; Lin, L.-K.; Zhang, J.-P.; Zhu, G.-J.; Huang, D.-W.; Zhang, S.-Y. (2007). "A New Species of Barbastella (Chiroptera: Vespertilionidae) from North China". Journal of Mammalogy. 88 (6): 1393–1403. doi:10.1644/07-MAMM-A-114R2.1.
  4. ↑ Esselstyn, J.A. (2007). "A New Species of Stripe-Faced Fruit Bat (Chiroptera: Pteropodidae: Styloctenium) from the Philippines" (PDF). Journal of Mammalogy. 88 (4): 951–958. doi:10.1644/06-mamm-a-294r.1.
  5. ↑ Bates, P.J.J.; Rossiter, S.J.; Suyanto, A.; Kingston, T. (2007). "A new species of Hipposideros (Chiroptera: Hipposideridae) from Sulawesi" (PDF). Acta Chiropterologica. 9 (1): 13–26. doi:10.3161/1733-5329(2007)9[13:ansohc]2.0.co;2. Archived from the original (PDF) on 2011-07-14.
  6. ↑ Francis, C.M.; Kingston, T.; Zubaid, A. (2007). "A new species of Kerivoula (Chiroptera: Vespertilionidae) from peninsular Malaysia" (PDF). Acta Chiropterologica. 9 (1): 1–12. doi:10.3161/1733-5329(2007)9[1:ansokc]2.0.co;2.
  7. ↑ João M. D. Miranda, Itiberê P. Bernardi & Fernando C. Passos (2006). "A new species of Eptesicus (Mammalia: Chiroptera: Vespertilionidae) from the Atlantic Forest, Brazil" (PDF). Zootaxa. 1383: 57–68.
  8. ↑ Woodman, N.; Timm, R.M. (2006). "Characters and phylogenetic relationships of nectar-feeding bats, with descriptions of new Lonchophylla from western South America (Mammalia: Chiroptera: Phyllostomidae: Lonchophyllini)" (PDF). Proceedings of the Biological Society of Washington. 119 (4): 437–476. doi:10.2988/0006-324x(2006)119[437:capron]2.0.co;2.
  9. ↑ Thabah, A.; Rossiter, S.J.; Kingston, T.; Zhang, S.; Parsons, S.; Mya Mya, Khin; Zubaid, A.; Jones, G. (2006). "Genetic divergence and echolocation call frequency in cryptic species of Hipposideros larvatus s.l. (Chiroptera: Hipposideridae) from the Indo-Malayan region" (PDF). Biological Journal of the Linnean Society. 88: 119–130. doi:10.1111/j.1095-8312.2006.00602.x. Archived from the original (PDF) on 2011-07-14.
  10. ↑ Guillén-Servent, A. & C.M. Francis (2006). "A new species of bat of the Hipposideros bicolor group (Chiroptera: Hipposideridae) from Central Laos, with evidence of convergent evolution with Sudaic taxa". Acta Chiropterologica. 8 (1): 39–61. doi:10.3161/1733-5329(2006)8[39:ANSOBO]2.0.CO;2.
  11. ↑ Goodman, S.M., F.H. Ratrimomanarivo, and F.H. Randrianadrianina (2006). "A new species of Scotophilus (Chiroptera: Vespertilionidae) from western Madagascar". Acta Chiropterologica. 8 (1): 21–37. doi:10.3161/1733-5329(2006)8[21:ANSOSC]2.0.CO;2.
  12. ↑ Kuo, H.-C.; Fang, Y.-P.; Csorba, G.; Lee, L.-L. (2006). "The definition of Harpiola (Vespertilionidae: Murininae) and the description of a new species from Taiwan" (PDF). Acta Chiropterologica. 8 (1): 11–19. doi:10.3161/1733-5329(2006)8[11:tdohvm]2.0.co;2.
  13. ↑ Spitzenberger, F.; Strelkov, P.P.; Winkler, H.; Haring, E. (2006). "A preliminary revision of the genus Plecotus (Chiroptera: Vespertilionidae) based on genetic and morphological results" (PDF). Zoologica Scripta. 35 (3): 187–230. doi:10.1111/j.1463-6409.2006.00224.x.
  14. ↑ Solari, S.; Baker, R.J. (2006). "Mitochondrial DNA Sequence, Karyotypic, and Morphological Variation in the Carollia castanea Species Complex (Chiroptera: Phyllostomidae) with Description of a New Species" (PDF). Occasional Papers of the Museum of Texas Tech University. 254: 1–16.
  15. ↑ Muchhala, N.; Mena, V.; Albuja, V. (2005). "A New Species of Anoura (Chiroptera: Phyllostomidae) from the Ecuadorian Andes" (PDF). Journal of Mammalogy. 86 (3): 457–461. doi:10.1644/1545-1542(2005)86[457:ansoac]2.0.co;2.
  16. ↑ Gregorin, R.; Ditchfield, A.D. (2005). "New Genus and Species of Nectar-Feeding Bat in the Tribe Lonchophyllini (Phyllostomidae: Glossophaginae) from Northeastern Brazil". Journal of Mammalogy. 86 (2): 403–414. doi:10.1644/BRB-229.1.
  17. ↑ Helgen, K.M. (2005). "Systematics of the Pacific monkey-faced bats (Chiroptera: Pteropodidae), with a new species of Pteralopex and a new Fijian genus". Systematics and Biodiversity. 3 (4): 433–453. doi:10.1017/S1477200005001702.
  18. ↑ Yoshiyuki, M.; Lim, B.L. (2005). "A New Horseshoe Bat, Rhinolophus chiewkweeae (Chiroptera, Rhinolophidae), from Malaysia" (PDF). Bulletin of the National Science Museum, Tokyo, Series A, Zoology. 31 (1): 29–36.
  19. ↑ Albuja, V; Gardner, A.L.; Graves, Gary R. (2005). Graves, Gary R., ed. "A new species of Lonchophylla Thomas (Chiroptera: Phyllostomidae) from Ecuador". Proceedings of the Biological Society of Washington. 118 (2): 442–449. doi:10.2988/0006-324X(2005)118[442:ANSOLT]2.0.CO;2.
  20. ↑ Happold, M. (2005). "A new species of Myotis (Chiroptera:Vespertilionidae) from central Africa". Acta Chiropterologica. 7 (1): 9–21. doi:10.3161/1733-5329(2005)7[9:ANSOMC]2.0.CO;2.
  21. ↑ Csorba, G. & P.J.J. Bates (2005). "Description of a new species of Murina from Cambodia (Chiroptera: Vespertilionidae: Murininae". Acta Chiropterologica. 7 (1): 1–7. doi:10.3161/1733-5329(2005)7[1:DOANSO]2.0.CO;2.
  22. ↑ Goodman, S.M.; Jenkins, R.K.B.; Ratrimomanarivo, F.H. (2005). "A review of the genus Scotophilus (Mammalia, Chiroptera, Vespertilionidae) on Madagascar, with the description of a new species" (PDF). Zoosystema. 27 (4): 867–882. Archived from the original (PDF) on 2006-11-27.
  23. ↑ Bergmans, W. & N.J. van Strien (2004). "Systematic notes on a collection of bats from Malawi. I. Megachiroptera: Epomophorinae and Rousettinae (Mammalia, Chiroptera)". Acta Chiropterologica. 6 (2): 249–268. doi:10.3161/001.006.0205.
  24. ↑ Goodman, S.M. & S.G. Cardiff (2004). "A new species of Chaerephon (Molossidae) from Madagascar with notes on other members of the family". Acta Chiropterologica. 6 (2): 227–248. doi:10.3161/001.006.0204.
  25. ↑ Bates, P.J.J.; Struebig, M.J.; Rossiter, S.J.; Kingston, T.; Sai; Lin Oo, Sein; Mya Mya, Khin (2004). "A new species of Kerivoula (Chiroptera: Vespertilionidae) from Myanmar (Burma)" (PDF). Acta Chiropterologica. 6 (2): 219–226. doi:10.3161/001.006.0203. Archived from the original (PDF) on 2011-07-25.
  26. ↑ Benda, P., P. Hulva, and J. Gaisler (2004). "Systematic status of African populations of Pipistrellus pipistrellus complex (Chiroptera: Vespertilionidae), with a description of a new species from Cyrenaica, Libya". Acta Chiropterologica. 6 (2): 193–217. doi:10.3161/001.006.0202.
  27. ↑ Dávalos, L.M. (2004). "A New Chocoan Species of Lonchophylla (Chiroptera: Phyllostomidae)" (PDF). American Museum Novitates. 3426: 1–14. doi:10.1206/0003-0082(2004)426<0001:ancsol>2.0.co;2.
  28. ↑ Fonseca, R.M.; Pinto, C.M. (2004). "A New Lophostoma (Chiroptera: Phyllostomidae: Phyllostominae) from the Amazonia of Ecuador" (PDF). Occasional Papers of the Museum of Texas Tech University. 242: 1–9.
  29. ↑ Baker, R.J.; Fonseca, R.M.; Parish, D.A.; Phillips, C.J.; Hoffmann, F.G. (2004). "New Bat of the Genus Lophostoma (Phyllostomidae: Phyllostominae) from Northwestern Ecuador" (PDF). Occasional Papers of the Museum of Texas Tech University. 232: 1–16.
  30. ↑ Pacheco, V.; Solari, S.; Velazco, P.M. (2004). "A New Species of Carollia (Chiroptera: Phyllostomidae) from the Andes of Peru and Bolivia" (PDF). Occasional Papers of the Museum of Texas Tech University. 236: 1–15.
  31. ↑ Maryanto, I.; Yani, M. (2003). "A new species of Rousettus (Chiroptera: Pteropodidae) from Lore Lindu, Central Sulawesi". Mammal Study. 28: 111–120. doi:10.3106/mammalstudy.28.111.
  32. ↑ Robinson, M.F., P.D. Jenkins, C.M. Francis, and A.J.C. Fulford (2003). "A new species of the Hipposideros pratti group (Chiroptera, Hipposideridae) from Lao PDR and Vietnam". Acta Chiropterologica. 5 (1): 31–48. doi:10.3161/001.005.0103.
  33. ↑ Parnaby, H.E. (2002). "A new species of long-eared bat (Nyctophilus: Vespertilionidae) from New Caledonia" (PDF). Australian Mammalogy. 23: 115–124. doi:10.1071/am01115. Archived from the original (PDF) on 2008-08-23.
  34. ↑ Baker, R.J.; Solari, S.; Hoffmann, F.G. (2002). "A New Central American Species from the Carollia brevicauda Complex" (PDF). Occasional Papers of the Museum of Texas Tech University. 217: 1–12.
  35. ↑ Simmons, N.B.; Voss, R.S.; Fleck, D.W. (2002). "A New Amazonian Species of Micronycteris (Chiroptera: Phyllostomidae) with Notes on the Roosting Behavior of Sympatric Congeners" (PDF). American Museum Novitates. 3358: 1–14. doi:10.1206/0003-0082(2002)358<0001:anasom>2.0.co;2.
  36. ↑ Cotterill, F.P.D. (2002). "A new species of horseshoe bat (Microchiroptera: Rhinolophidae) from south-central Africa: with comments on its affinities and evolution, and the characterization of rhinolophid species". Journal of Zoology. 256 (2): 165–179. doi:10.1017/S0952836902000201.
  37. ↑ Parnaby, H.E. (2002). "A taxonomic review of the genus Pteralopex (Chiroptera: Pteropodidae), the monkey-faced bats of the south-western Pacific" (PDF). Australian Mammalogy. 23: 145–162. doi:10.1071/am01145. Archived from the original (PDF) on 2008-08-23.
  38. ↑ Mucedda, M., A. Kiefer, E. Pidinchedda, and M. Veith (2002). "A new species of long-eared bat (Chiroptera, Vespertilionidae) from Sardinia". Acta Chiropterologica. 4 (2): 121–135. doi:10.3161/001.004.0202.
  39. ↑ Fahr, J.; Vierhaus, H.; Hutterer, R.; Kock, D. (2002). "A revision of the Rhinolophus maclaudi species group with the description of a new species from West Africa (Chiroptera: Rhinolophidae)" (PDF). Myotis. 40: 95–126. Archived from the original (PDF) on 2005-09-22.
  40. ↑ Richards, G.C. & L.S. Hall (2002). "A new flying-fox of the genus Pteropus (Chiroptera: Pteropodidae) from Torres Strait, Australia". Australian Zoologist (32): 69–75.
  41. ↑ Helgen, K.M. (2004). "On the identity of flying-foxes, genus Pteropus (Mammalia: Chiroptera), from islands in the Torres Strait, Australia". Zootaxa (780): 1–14.
  42. ↑ Kruskop, S.V.; Tsytsulina, K.A. (2001). "A new big-footed mouse-eared bat Myotis annamiticus sp. nov. (Vespertilionidae, Chiroptera) from Vietnam" (PDF). Mammalia. 65 (1): 63–72. doi:10.1515/mamm.2001.65.1.63.
  43. ↑ Von Helversen, O.; Heller, K.-G.; Mayer, F.; Nemeth, A.; Volleth, M.; Gombkötö, P. (2001). "Cryptic mammalian species: a new species of whiskered bat (Myotis alcathoe n. sp.) in Europe" (PDF). Naturwissenschaften. 88: 217–223. doi:10.1007/s001140100225. PMID 11482435.
  44. ↑ Cuartas, C.A.; Muñoz, J.; González, M. (2001). "Una nueva especie de Carollia Gray, 1838 (Chiroptera: Phyllostomidae) de Colombia". Actualidades Biológicas. 23 (75): 63–73.
  45. ↑ Muñoz, J.; Cuartas, C.A. (2001). "Saccopteryx antioquensis n. sp. (Chiroptera: Emballonuridae) del noroeste de Colombia". Actualidades Biológicas. 23 (75): 53–61.
  46. ↑ Bergmans, W. (2001). "Notes on distribution and taxonomy of Australasian bats. I. Pteropodinae and Nyctimeninae (Mammalia, Megachiroptera, Pteropodidae)". Beaufortia Bulletin Zoological Museum (51 (8)): 119–152.
  47. ↑ Eger, J.L. & D.A. Schlitter (2001). "A new species of Glauconycteris from West Africa (Chiroptera: Vespertilionidae)". Acta Chiropterologica. 3 (1): 1–10.
  48. ↑ Eger, J.L. (2001). "Emendation of Glauconycteris curryi". Acta Chiropterologica. 3 (2): 248.
  49. ↑ Kruskop, S.V.; Lavrenchenko, L.A. (2000). "A new species of long-eared bat (Plecotus; Vespertilionidae, Mammalia) from Ethiopia" (PDF). Myotis. 38: 5–17.
  50. ↑ Contreras, M.; Cadena, A. (2000). "Una nueva especie del género Sturnira (Chiroptera, Phyllostomidae) de los Andes colombianos" (PDF). Revista de la Academia Colombiana de Ciencias Exactas, Físicas y Naturales. 24 (91): 285–287. Archived from the original (PDF) on 2008-11-20.
  51. ↑ Kock, D.; Csorba, G.; Howell, K.M. (2000). "Rhinolophus maendeleo n. sp. from Tanzania, a horseshoe bat noteworthy for its systematics and biogeography (Mammalia, Chiroptera, Rhinolophidae)". Senckenbergiana Biologica. 80: 233–239.
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